The thread logic of centre-lock wheels depended on the conical load-bearing surfaces of hub and wheel - when the thing is slack the hub drops a few thou causing the hub and wheel cones to touch (and bear the load) where they have slightly different diameters. What is then required is that the rotation with those dissimilar diameters drags the nut in the tightening sense, for which one needs handed threads.
The conventional nut had a female cone locating on the wheel's male cone which gives the easily-remembered "right-hand thread on the left-hand side/left-hand thread on the right-hand side" logic. Lotus confused many folk with the Elan which had the female cone on the wheel so the left was left and the right right.
What has been proven time and again is that if someone fits them swapped over the wheels do stay on if the nuts are very tight but come off the moment there's a bit of slack.to unwind the nut.
In racing cars that may have all changed by now, but not in 1967, I'd say and as the OP has seen the "left-hand thread on right-hand side" it all hangs together.
Edited by Allan Lupton, 31 March 2020 - 11:48.